Anthropogenic Drivers and Ecological Consequences of Global Insect Population Attrition

The observation that vehicle windshields remain noticeably cleaner during long-distance travel compared to previous decades has transitioned from a common anecdote to a central focal point of entomological concern. This "windshield phenomenon" serves as a visible, albeit qualitative, proxy for a much deeper and more pervasive ecological crisis: the rapid decline of flying insect biomass across diverse terrestrial landscapes.1 While early skepticism questioned whether such observations were merely artifacts of human memory or improvements in vehicle aerodynamics, longitudinal scientific investigations have since provided robust quantitative support for these anecdotal claims.4 Research conducted over the past several decades reveals that the decline is not isolated to specific charismatic species but reflects a systemic reduction in the total biomass and abundance of the entomofauna, with profound implications for the stability of global food webs and the continued provision of essential ecosystem services.6

Quantitative Analysis of the Windshield Phenomenon as a Biological Proxy

The transformation of the windshield phenomenon from a social observation into a scientific metric required rigorous longitudinal monitoring and cross-validation with established entomological sampling methods. In Denmark, a 20-year study (1997–2017) utilized car windscreens as a novel quantitative tool to measure insect abundance along specific road transects.4 By driving at a standardized speed of  and controlling for environmental variables such as temperature, wind speed, and time of day, researchers documented a dramatic reduction in insect "splatter".2 The data indicated biomass reductions of 80% and 97% across two distinct transects, demonstrating that the observed decline was both geographically widespread and statistically significant even after accounting for climatic fluctuations.4

Crucially, these windscreen-based results were cross-validated using traditional methods, including sweep-net sampling and sticky plate traps. The high degree of positive correlation between the number of insects killed on car windscreens and the abundance of insects captured in nets or on plates suggests that the windshield phenomenon is a biologically relevant and accurate reflection of local insect availability.4 Similar "splatometer" surveys in the United Kingdom supported these findings; a study by the Kent Wildlife Trust reported a 72% decrease in insect collisions on vehicle number plates between 2004 and 2021.2

Technical assessments of the windshield phenomenon must also account for changes in automotive design. It has been hypothesized that modern, aerodynamic vehicle shapes might push insects over the car rather than resulting in a collision. However, empirical tests suggest the opposite: modern vehicles, with their more aerodynamic body shapes, often kill more insects than boxier vintage models because the latter push a larger volume of air—and thus insects—away from the surface.2 This aerodynamic nuance reinforces the gravity of the observed decline; if modern cars are more efficient at collecting insects, the reduction in visible splatter suggests an even more precipitous drop in the underlying population density.2

Comparative Findings of Windshield and Collision Surveys

Global Patterns of Biomass and Abundance Attrition

The most alarming evidence of insect decline comes from long-term biomass monitoring using Malaise traps. The Krefeld Entomological Society’s 27-year study in Germany remains a cornerstone of this research, documenting a seasonal decline of 76% and a mid-summer peak decline of 82% in total flying insect biomass within 63 nature protection areas.7 Because these declines occurred within regions supposedly protected from direct habitat destruction, the results imply that large-scale, diffuse factors—such as pesticide drift or climate change—are operating across the landscape regardless of local conservation status.7

The German data is reflective of a broader trend observed across Northern Europe. In Sweden’s Kullaberg nature reserve, researchers found that more than a quarter of the 600 species of butterflies and moths once present had been lost over a 50-year period.11 Similarly, leafhoppers and planthoppers, which constitute a significant portion of the biomass in European grasslands, saw their abundance in Germany plunge by 66% between 1960 and 2010.11 These findings suggest that the decline is not restricted to specific lineages but represents a systemic thinning of the insect community across multiple orders.6

In North America, the trends are equally troubling. A study conducted in the Colorado mountains between 2004 and 2024 documented a 6.6% annual decline in flying insect density in a remote meadow, totaling a 72.4% drop over 20 years.12 The remote nature of this study site is particularly significant, as it suggests that insect populations are collapsing even in areas free from direct human interference or intensive agriculture, pointing toward climate change as a pervasive, global driver.12

The Controversial Case of Tropical Forest Declines

The application of high-latitude findings to tropical regions has been a subject of intense scientific debate. A 2018 report by Lister and Garcia claimed a "catastrophic collapse" in the Luquillo Experimental Forest in Puerto Rico, asserting that insect biomass had fallen by a factor of 10 to 60 since the 1970s.5 The study linked this decline to a  increase in mean maximum temperatures, suggesting that tropical insects, having evolved in stable climates, are nearing their physiological thermal limits.5

However, this narrative has been contested by other researchers who argue that the Lister and Garcia data suffers from unjustified assumptions and analytical issues.15 Critics point out that the sampling periods coincided with major forest disturbances; specifically, the initial 1970s samples were taken during a period of forest recovery following Hurricane David, while later samples were influenced by the long-term succession following Hurricanes Hugo and Maria.16 Subsequent analyses suggest that canopy arthropod populations in Luquillo were just as likely to increase as they were to decrease in response to temperature shifts, and that hurricane-induced successional dynamics are the primary drivers of population variance.16 This controversy highlights the complexity of disentangling climate signals from natural disturbance cycles, particularly in the tropics.18

Longitudinal Biomass and Abundance Trends by Region

Anthropogenic Drivers of Population Collapse

The causes of insect decline are multifaceted, often characterized as a "death by a thousand cuts" where multiple stressors interact synergistically to undermine population resilience.15 While local factors vary, the primary drivers consistently identified across global meta-analyses include agricultural intensification, the ubiquitous use of synthetic pesticides, habitat fragmentation, and the escalating effects of climate change.6

Agricultural Intensification and Habitat Loss

The conversion of diverse natural landscapes into large-scale monocultures represents perhaps the most significant historical driver of insect loss. Intensive agriculture eliminates the "structural complexity" of the environment, removing hedgerows, forest edges, and fallow fields that once served as critical nesting sites and floral resources for diverse insect communities.1 This process creates "biological deserts" where only a handful of specialized or pest species can survive, leading to a homogenization of the entomofauna where specialized species are replaced by a few resilient generalists.10

Furthermore, agricultural intensification is often accompanied by the heavy use of synthetic fertilizers, which leads to soil acidification and eutrophication of nearby water bodies.6 These chemical changes directly impact the nutritional quality of host plants and the survival of aquatic insect larvae.8 For example, the 66% plunge in leafhopper abundance in German grasslands has been largely attributed to soil acidification resulting from decades of nitrogen application, which fundamentally altered the chemistry of the grasses on which they feed.11

The Role of Neurotoxic Pesticides

The introduction and widespread adoption of new classes of systemic insecticides, particularly neonicotinoids and fipronil, have correlated with the most precipitous declines observed in the last two decades.11 Unlike traditional contact pesticides, neonicotinoids are absorbed into plant tissues, making the entire plant—including nectar and pollen—toxic to insects.8 These chemicals persist in the environment for years and are highly mobile in water, leading to chronic, sublethal exposure for non-target species such as wild bees, butterflies, and aquatic invertebrates.8

At the physiological level, exposure to these compounds impairs an insect’s ability to navigate, forage, and reproduce. Studies have shown that even at concentrations several orders of magnitude lower than those used in legacy pesticides like DDT, neonicotinoids interfere with neurobiological functions, leading to immunosuppression and increased susceptibility to pathogens.8 This creates a "pesticide treadmill" where the loss of beneficial predatory insects necessitates even higher applications of chemicals to manage resurgent pest populations, further decimating the ecological infrastructure that naturally regulates agriculture.25

Climate Change: Thermal Physiology and Hypoxia

Insects are ectothermic organisms, meaning their physiological processes, activity levels, and reproductive rates are inextricably linked to ambient temperatures.28 As global temperatures rise and weather patterns become more erratic, many species are being pushed beyond their thermal optima.3 In temperate regions, species may shift their ranges poleward or to higher elevations as things get warmer, which can result in localized declines even if the global population remains stable.3 However, in mountain or island environments, there is often nowhere to migrate, leading to "escalator to extinction" scenarios.30

Recent research has also identified a link between pesticide exposure and climate stress. Neonicotinoids and other systemic chemicals impair mitochondrial respiration, creating a hypoxia-like stress response in insects.8 This respiratory inhibition reduces their tolerance to rising temperatures, as oxygen demands increase with metabolic rates.8 Additionally, increased atmospheric  levels promote faster plant growth but result in biomass that is lower in essential nutrients, effectively starving herbivorous insects even when food is abundant.6

Secondary Drivers: Light, Noise, and Invasive Species

An often-overlooked factor in the decline of flying insects is the rapid expansion of artificial light at night (ALAN). Light pollution disorients nocturnal fliers such as moths and fireflies, disrupting their mating behaviors and increasing their vulnerability to predation.21 For many species, artificial light acts as a "population sink," drawing insects away from suitable habitats and toward urban areas where they perish from exhaustion or predation.3

Biological factors also play a critical role. Invasive species introduce competition and novel pathogens that native insects are ill-equipped to handle. For instance, the Asian Lady Beetle contains fungal parasites that are lethal to native ladybug species.21 Global trade has also facilitated the spread of pathogens from managed honey bees to wild bumble bee populations, contributing to the decline of the latter.21

Detriments to Ecosystem Stability and Trophic Cascades

The precipitous decline in insect biomass is not merely a loss of biodiversity; it is a structural failure of the foundations of most terrestrial and freshwater ecosystems. Insects represent approximately 80% of all animal species and are the primary conduit for energy transfer from plants to higher trophic levels.6

Bottom-Up Trophic Cascades

The most direct consequence of insect decline is the subsequent collapse of populations that rely on them for food. Insectivorous birds, reptiles, amphibians, and bats are experiencing significant population drops that correlate closely with insect biomass trends.10 In Denmark, the decline in insects killed on windscreens was found to be a strong predictor of the rate at which barn swallows (Hirundo rustica) could feed their nestlings.4 As food availability decreased, the size of the breeding populations of aerially insectivorous birds fell in parallel, illustrating a clear bottom-up trophic link.4

In the Luquillo Forest, the initial (though contested) report of insect loss was accompanied by synchronous declines in the lizards and frogs that consume them.5 Even if the exact cause of these declines remains a subject of scientific debate, the consensus is that the removal of such a large portion of the food web’s base will inevitably lead to a simplification of the entire ecosystem, reducing its resilience to further disturbances.14 The Puerto Rican tody (Todus mexicanus), a bird that feeds almost exclusively on insects, was reported to have dropped by 90% in some surveys, though other data suggests capture rates have remained more stable, illustrating the challenges of assessing trophic collapse.14

Disruption of Essential Ecosystem Services

The services provided by insects are often undervalued because they are performed invisibly and without direct cost to human society. However, the economic and ecological costs of replacing these services are staggering. Insects are responsible for four primary categories of ecosystem functioning:

1. Pollination: Approximately 80% of wild plants and 75% of global food crops rely on animal-mediated pollination.27
2. Nutrient Cycling and Decomposition: Decomposers break down organic matter, fallen leaves, and animal carcasses, recycling nutrients like nitrogen and phosphorus back into the soil for plant growth.26
3. Natural Pest Control: Beneficial predatory insects, such as ladybugs and parasitic wasps, regulate the populations of herbivorous insects that would otherwise devastate crops, saving billions in chemical control costs.25
4. Soil Aeration and Engineering: Soil insects like ants and termites move vast quantities of earth, influencing soil structure, drainage, and fertility.34

The valuation of these services in the United States alone is estimated at approximately  annually.32 Globally, the loss of pollination services is projected to result in a welfare loss of up to .36

Extension of the Decline to Non-Flying Insects

While the "windshield phenomenon" specifically highlights the decline of flying insects, current research indicates that the trend extends to many non-flying and ground-dwelling taxa, although these groups are often less extensively monitored.1 Soil macrofauna, including ants, beetles, and termites, are critical engineers of the subterranean world, yet they face many of the same anthropogenic pressures as their flying counterparts.35

Trends in Soil and Ground-Dwelling Arthropods

Comprehensive assessments of terrestrial arthropods suggest that while flying insects may show more immediate biomass drops in Malaise traps, ground-dwelling species are also in retreat.38 In Puerto Rico, canopy-dwelling insects fell by 98%, while ground-foraging insects fell by 78% over the same 35-year study period.14 This suggests that while flying insects may be more sensitive to rapid environmental shifts due to their high metabolic costs and exposure to aerial pollutants, ground-dwelling insects are equally vulnerable to habitat degradation and soil contamination.39

Ground beetles (Carabidae), which are predominantly non-flying or reluctant fliers, have shown dramatic declines in the United Kingdom, where nearly three-quarters of the 68 species studied have decreased in abundance.11 In North American tallgrass prairies, grasshopper populations have declined due to changes in burning frequency and plant nutrient quality.24 Similarly, aquatic insects in their larval (non-flying) stages are highly sensitive to synthetic fertilizers and pesticides that leach into water bodies, leading to "considerable losses" in dragonflies and caddisflies.8

Soil Macrofauna and Global Biomass Distribution

The decline of non-flying insects is particularly concerning given their role in maintaining soil health. Ants and termites alone contribute significantly to global biomass; termites represent approximately 40% of the soil arthropod biomass, while ants contribute around 10%.38 Ants constructed extensive nests that move enough soil to cover fields with a  layer annually in some regions, a process vital for soil aeration.35

Biological invasions represent a unique threat to non-flying insects. In North American forests, the introduction of invasive earthworms has been shown to radically alter soil structure and leaf litter layers, leading to a significant decrease in the total arthropod abundance, biomass, and species richness.24 This homogenization of the soil community mirrors the trends seen above-ground, as specialized native species are displaced by invasive generalists.24

Distribution of Global Terrestrial Arthropod Biomass

Economic Detriments and Risks to Global Food Security

The potential for a "pollinator collapse" represents one of the most severe economic risks associated with insect decline. While the total loss of all pollinators is an extreme scenario, even partial declines are already impacting agricultural yields and human nutrition.8

Yield Stability and Nutrient Availability

Research models suggest that a collapse of wild pollinators would lead to an average 8% decline in European crop yields, with much higher losses for nutrient-dense crops such as fruits, vegetables, and oilseeds.40 In higher-income countries, the impact is often masked by the use of managed honeybees (Apis mellifera), yet managed bees cannot fully replace the efficiency or diversity of wild pollinators.15 In North America, nearly one in four native bee species is at risk of extinction, and 19% of butterfly species are similarly threatened.21

The consequences for human health are profound. Pollinator-dependent crops provide the majority of the world’s Vitamin A, folate, and essential fatty acids. A significant reduction in pollinator populations could lead to widespread micronutrient deficiencies, particularly in developing nations where diets are less diversified.25 Global trade complicates this issue; while about 17% of global crop production value depends on pollination, these crops make up 28% of global agricultural trade, meaning that declines in one region can trigger price spikes and nutritional shortages worldwide.36

Disproportionate Impact on Developing Nations

Smallholder farmers in low- and middle-income countries are the most vulnerable to insect loss. Many high-value "cash crops" that drive the economies of these regions—such as coffee, cocoa, almonds, and avocados—are highly dependent on animal pollination.25 In these contexts, the loss of natural pest control and pollination services cannot be easily mitigated by expensive technological interventions or chemical substitutes.25 This exacerbates global inequality, as the economic shock of pollinator decline hits the world’s poorest farmers the hardest while the environmental costs of the pesticides that drive the decline are not reflected in their market prices.25

Economic Valuation of Insect-Mediated Services

Scientific Discourse, Critiques, and the Path Forward

The "insect apocalypse" narrative, while powerful for mobilizing public concern, has been criticized for overextrapolation. Some entomologists note that applying localized data to the global scale is not always statistically sound, and that some species are actually increasing in abundance or shifting their ranges poleward.3 For example, temperate insects limited by winter temperatures may thrive as global temperatures rise, and anthropophilic taxa like cockroaches and houseflies are likely to boom as biodiversity elsewhere declines.15

Despite these nuances, the consensus across 175 scientific reviews and thousands of site evaluations is that the rate of insect decline is staggering—estimated at  annually, or eight times faster than that of mammals or birds.8 This signals a fundamental breakdown in ecological stability that requires urgent policy intervention.6

Policy Interventions and Conservation Strategies

Recognizing the severity of the decline, several international bodies have begun implementing legislative frameworks. The European Union has developed the EU Pollinators Initiative and the Nature Restoration Law (NRL), which aim to restore ecosystems across 20% of the EU's land area by 2030.42 Article 10 of the NRL establishes a binding target for Member States to reverse the decline of pollinators and achieve an increasing trend thereafter, measured every six years.42

However, critics argue that these policies often fail to address the root causes of destruction, such as intensive livestock farming and the overuse of pesticides.48 Effective conservation requires a multi-pronged approach:

• Habitat Restoration: Protecting native vegetation, wetlands, and forest edges to create safe havens for diverse communities.26
• Pesticide Reform: Phasing out neonicotinoids and glyphosate while promoting organic and third-party certified farming products.21
• Urban Integration: Reducing grass lawns and increasing native plant richness in developed areas to support pollinators.21
• Global Monitoring: Expanding long-term insect data collection beyond Europe and North America into the tropics and Asia to close critical knowledge gaps.19

The decline of flying and non-flying insects represents an unprecedented threat to the biophysical systems that support human life. The "windshield phenomenon" is not merely a nostalgic observation of the past but a stark warning about the future of global biodiversity and food security.1 Halting this trend will require a coordinated global effort to mitigate the "death by a thousand cuts" and restore the intricate web of life that insects maintain.15

Works cited

1. The “Windshield Phenomenon” – Where Have All the Insects Gone?, accessed February 5, 2026, https://www.irejn.org/the-windshield-phenomenon-where-have-all-the-insects-gone/
2. Windshield phenomenon - Wikipedia, accessed February 5, 2026, https://en.wikipedia.org/wiki/Windshield_phenomenon
3. When insects hit the windshield, science starts talking - AGDAILY, accessed February 5, 2026, https://www.agdaily.com/insights/when-insects-hit-the-windshield-science-starts-talking/
4. Parallel declines in abundance of insects and insectivorous birds in ..., accessed February 5, 2026, https://pmc.ncbi.nlm.nih.gov/articles/PMC6580276/
5. What Is the 'Windshield Phenomenon'? - Treehugger, accessed February 5, 2026, https://www.treehugger.com/what-is-the-windshield-phenomenon-4862086
6. Decline in insect populations - Wikipedia, accessed February 5, 2026, https://en.wikipedia.org/wiki/Decline_in_insect_populations
7. More than 75 percent decline over 27 years in total flying insect ..., accessed February 5, 2026, https://pmc.ncbi.nlm.nih.gov/articles/PMC5646769/
8. Insect Decline in the Anthropocene: Historical Parallels and Emerging Monitoring Tools, accessed February 5, 2026, https://www.mdpi.com/2075-4450/16/8/841
9. More Than 75 Percent Decrease in Total Flying Insect Biomass Over 27 Years, accessed February 5, 2026, https://www.dadant.com/75-percent-decrease-total-flying-insect-biomass-27-years/
10. 'Half the tree of life': ecologists' horror as nature reserves are emptied of insects - The Guardian, accessed February 5, 2026, https://www.theguardian.com/environment/2025/jun/03/climate-species-collapse-ecology-insects-nature-reserves-aoe
11. Plummeting insect numbers 'threaten collapse of nature' | Insects ..., accessed February 5, 2026, https://www.theguardian.com/environment/2019/feb/10/plummeting-insect-numbers-threaten-collapse-of-nature
12. Insect populations drop even without direct human interference, a new study finds - WHRO, accessed February 5, 2026, https://www.whro.org/2025-09-12/insect-populations-drop-even-without-direct-human-interference-a-new-study-finds
13. Climate-driven declines in arthropod abundance restructure a rainforest food web | PNAS, accessed February 5, 2026, https://www.pnas.org/doi/10.1073/pnas.1722477115
14. Insect collapse: 'We are destroying our life support systems' - The Guardian, accessed February 5, 2026, https://www.theguardian.com/environment/2019/jan/15/insect-collapse-we-are-destroying-our-life-support-systems
15. Insect decline in the Anthropocene: Death by a thousand cuts - PNAS, accessed February 5, 2026, https://www.pnas.org/doi/10.1073/pnas.2023989118
16. Arthropods are not declining but are responsive to disturbance in the Luquillo Experimental Forest, Puerto Rico | PNAS, accessed February 5, 2026, https://www.pnas.org/doi/10.1073/pnas.2002556117
17. Populations are not declining and food webs are not collapsing at the Luquillo Experimental Forest - PMC - NIH, accessed February 5, 2026, https://pmc.ncbi.nlm.nih.gov/articles/PMC6589750/
18. Study explores decline of insect populations in tropical forests - Griffith News, accessed February 5, 2026, https://news.griffith.edu.au/2025/04/08/study-explores-decline-of-insect-populations-in-tropical-forests/
19. Declining insect biodiversity in the tropics - ScienceDaily, accessed February 5, 2026, https://www.sciencedaily.com/releases/2025/04/250408121928.htm
20. Insect Research - Catawba College, accessed February 5, 2026, https://catawba.edu/news/all-news/2023/insect-research/
21. 5 Major Causes of Global Insect Declines & How to be Part of the Solut - Crown Bees, accessed February 5, 2026, https://crownbees.com/blogs/news/5-major-causes-of-global-insect-declines-how-to-be-part-of-the-solution
22. Insect declines and why they matter - Somerset Wildlife Trust, accessed February 5, 2026, https://www.somersetwildlife.org/sites/default/files/2019-11/FULL%20AFI%20REPORT%20WEB1_1.pdf
23. What Shrinking Insect Populations Mean for the Planet | Earth.Org, accessed February 5, 2026, https://earth.org/what-shrinking-insect-populations-mean-for-the-planet/
24. Less overall, but more of the same: drivers of insect population ... - NIH, accessed February 5, 2026, https://pmc.ncbi.nlm.nih.gov/articles/PMC10050920/
25. The threat declining insect populations pose to agriculture and ..., accessed February 5, 2026, https://www.ifpri.org/blog/threat-declining-insect-populations-pose-agriculture-and-development-and-what-we-can-do-about/
26. The global decline of insects - European Wilderness Society -, accessed February 5, 2026, https://wilderness-society.org/the-global-decline-of-insects/
27. Threats to Pollinators, accessed February 5, 2026, https://www.pollinator.org/threats
28. Insect Flight: State of the Field and Future Directions - PMC, accessed February 5, 2026, https://pmc.ncbi.nlm.nih.gov/articles/PMC11406162/
29. Life history evolution of insects in response to climate variation: seasonal timing versus thermal physiology | bioRxiv, accessed February 5, 2026, https://www.biorxiv.org/content/10.1101/2025.02.05.636251v1.full-text
30. Drop in insect populations not part of natural cycle - Charles University, accessed February 5, 2026, https://cuni.cz/UKEN-955.html
31. Wind plays a major but not exclusive role in the prevalence of insect flight loss on remote islands, accessed February 5, 2026, https://royalsocietypublishing.org/rspb/article/287/1940/20202121/85906/Wind-plays-a-major-but-not-exclusive-role-in-the
32. The Insect Effect: Insect Decline and the Future of Our Planet ..., accessed February 5, 2026, https://www.floridamuseum.ufl.edu/earth-systems/the-insect-effect/
33. How essential are pollinators for global food security? | World Economic Forum, accessed February 5, 2026, https://www.weforum.org/stories/2021/08/how-essential-are-pollinators-for-global-food-security/
34. SOIL MACROFAUNA FIELD MANUAL - Food and Agriculture Organization of the United Nations, accessed February 5, 2026, https://www.fao.org/4/i0211e/i0211e.pdf
35. Macrofauna » Insects » Ants, Bees and Wasps - SARE - Sustainable Agriculture Research and Education, accessed February 5, 2026, https://www.sare.org/publications/farming-with-soil-life/macrofauna-insects-ants-bees-and-wasps/
36. Pollinator declines, international trade and global food security, accessed February 5, 2026, https://eep.uni-hohenheim.de/uploads/media/1-s2.0-S0921800925000485-main.pdf
37. Knowledge on soil invertebrate macrofauna and bioturbating vertebrates: a global analysis using data science tools, accessed February 5, 2026, https://soil-organisms.org/SO/article/view/427
38. The global biomass and number of terrestrial arthropods - PMC - PubMed Central, accessed February 5, 2026, https://pmc.ncbi.nlm.nih.gov/articles/PMC9897674/
39. The Silent Extinction of Insects - National Biodiversity Data Centre, accessed February 5, 2026, https://biodiversityireland.ie/the-silent-extinction-of-insects/
40. The economic, agricultural, and food-security repercussions of a wild pollinator collapse in Europe, accessed February 5, 2026, https://www.fao.org/pollination/resources/knowledge/knowledge-product-detail/repercussions-of-a-wild-pollinator-collapse-europe/en
41. Economic Consequences of Pollinator Declines: A Synthesis Dana Marie Bauer and Ian Sue Wing Submitted ARER – August 1, 2010 - Boston University, accessed February 5, 2026, https://people.bu.edu/isw/papers/Bauer%20and%20SueWing%20ARER%20Final.pdf
42. Protecting and restoring Europe's wild pollinators and their habitats | Publications, accessed February 5, 2026, https://www.eea.europa.eu/en/analysis/publications/protecting-and-restoring-europes-wild-pollinators-and-their-habitats
43. Pollinator declines are an economic threat to global food systems - CentAUR, accessed February 5, 2026, https://centaur.reading.ac.uk/104396/1/People%20and%20Nature%20-%202022%20-%20Murphy%20-%20Globalisation%20and%20pollinators%20%20Pollinator%20declines%20are%20an%20economic%20threat%20to%20global.pdf
44. Insects are disappearing at an alarming rate worldwide. Insect populations had declined by 75% in less than three decades. The most cited driver for insect decline was agricultural intensification, via issues like land-use change and insecticides, with 500+ other interconnected drivers. : r/science - Reddit, accessed February 5, 2026, https://www.reddit.com/r/science/comments/1k5cduz/insects_are_disappearing_at_an_alarming_rate/
45. Nature Restoration Regulation - Environment - European Commission, accessed February 5, 2026, https://environment.ec.europa.eu/topics/nature-and-biodiversity/nature-restoration-regulation_en
46. Pollinators - Environment - European Commission, accessed February 5, 2026, https://environment.ec.europa.eu/topics/nature-and-biodiversity/pollinators_en
47. EU Pollinators Initiative (revised 2025) - Butterfly Conservation Europe, accessed February 5, 2026, https://www.bc-europe.eu/webpage.php?name=eu-pollinator-initiative
48. Assessing the effectiveness of the EU nature restoration law measured against the CBD target to preserve biodiversity - ResearchGate, accessed February 5, 2026, https://www.researchgate.net/publication/399957438_Assessing_the_effectiveness_of_the_EU_nature_restoration_law_measured_against_the_CBD_target_to_preserve_biodiversity